"therefore economies of scale should play apart in what they sell and how much they sell it for, and they should be much more competitive."
They do need to be more competitive but they are at a disadvantage in that they have to pay staff (however poor the staff are), upkeep of their stores and also pay business rates on each and every store. In other words they have very high overheads compared to the internet stores. Yes some of the stores you mention do have a high street presence but no where near the number of high street stores Jessops still has.
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